製 圖:莫妮卡.塞拉諾。ART: MONICA SERRANO AND CLARE TRAINOR, NGM STAFF. ANNIE ROTH; ALEXANDER STEGMAIER. SOURCES: MICHEL C. MILINKOVITCH AND OTHERS, NATURE, 2018;PAUL MANGER, UNIVERSITY OF THE WITWATERSRAND, JOHANNESBURG; IUCN/SSC AFRICAN ELEPHANT SPECIALIST GROUP